Just to draw reader's attention to one of the "Journal of Memetics" articles - "Do Animals Have Memes?".
Article link: http://cfpm.org/jom-emit/1999/vol3/reader_sm&laland_kn.html
It argues for the ubiquitous nature of memes among many animal species. It makes an excellent case that memetic transmission need not depend on imitation, and so that definitions of memes that invoke imitation are wrong.
The view the article criticises can be found here: Imitation and the definition of a meme.
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